Презентация на тему Аллопатрическое видообразование. Симпатрическое видообразование

B переводятся друг в друга масштабированием, а между группами

— масштабированием и срезающим преобразованием

от экспрессииBmp4. Bmp4 сильно коррелирует с масштабированием толщины клюва.

косвенных последствий отбора, другие — в результате самоорганизации

либо будут, либо не будет рогов. Изменения порога или

периода чувствительности к ювенильному гормону приводят к видовым различиям.

потери и приобретения полового диморфизма, а также изменения формы

рогов.

колонии, причём за 10-11 поколений большая часть хлорелл пребывала

в виде 8-клеточных колоний, а через 100 — в виде больших многоклеточных самовоспроизводящихся колоний.

Ничто не мешает сформировать колониальность/многоклеточность, если уже есть клеточная стенка/другие механизмы агрегации после митоза.

Ochromonas vallescia
(немного приукрашен)

Chlorella vulgaris

адаптивную ценность
Родопсины присутствуют у всех трёх доменов живых организмов.

Есть предположения, что изначально они не выполняли сенсорную функцию, а применялись для использования солнечной энергии.
В таком случае их можно рассматривать как преадаптацию к фотосенсорной функции

the Russian doll model, assumes that light sensitivity first

arose in cyanobacteria, the earliest known fossils on Earth. These cyanobacteria were subsequently taken up by eukaryotic red algae as primary chloroplasts surrounded by an outer and inner bacterial membrane separated by a proteoglycan layer.
Subsequently, the red algae were taken up by dinoflagellates as secondary chloroplasts surrounded by an additional third membrane coming from the primary red algal host. In some species of dinoflagellates like Erythropsis and Warnovia, which do not have any chloroplasts, these secondary chloroplasts may have been transformed into elaborate photoreceptor organelles as suggested by Greuet.
Indeed, the sequencing of the complete genome of the cyanobacterium Nostoc has revealed the presence of a proteorhodopsin gene (Kaneko et al. 2001). A proteorhodopsin gene has also been found in the dinoflagellate Pyrocystis and shown to be controlled by the internal clock (Okamoto and Hastings 2003)......»

Gehring, 2005

in eye evolution is the evolution of a light

receptor molecule which in all metazoans is rhodopsin. In the most ancestral metazoa, the sponges, a single Pax gene, but no opsin
gene has been found. In the cubozoan jellyfish Tripedalia, a unicellular photoreceptor has been described in the larva. The adult jellyfish has complex lens eyes that form under the control of PaxB, whereas the eyes of a hydrozoan jellyfish (Cladonema) are controlled by PaxA. We propose that from the unicellular
photoreceptor cell, the prototypic eye postulated by Darwin originated by a first step of cellular differentiation into a photoreceptor cell and a pigment cell, controlled by Pax6 and
MITF, respectively. (Gehring, 2012).

6 on the very top of the hierarchy. In

Drosophila, Pax 6 has undergone a gene duplication and twin of eyeless (toy) activates eyeless (ey) initiating the developmental pathway. Together these two genes activate the subordinate homeobox transcription factor sine oculis (so) by binding to five cis-regulatory elements identified in the so enhancer. It is interesting to note that two of the subfamilies of Six genes control eye development, whereas the third subfamily is involved in muscle development. This shows clearly that there are no functional constraints on transcription factors to control a particular developmental pathway. The So protein interacts with the protein encoded by eyes absent (eya) forming a hetero-dimer. Eya serves both as a co-activator in the nucleus and as a protein phosphatase in the cytoplasm. The transcription factors are connected by positive feed-back loops to dachshund (dac). There are two additional genes involved; one is optix which like so is member of the six gene family, and the other is the Pax gene eye gone (eyg) which apparently acts independently of ey and toy. In order to block eye development in Drosophila completely, all three Pax genes ey, toy, and eyg have to be mutated. Interestingly, the top of the cascade involves mostly positive regulation (activation) and positive feed-back loops.

genes in flies. A: Ectopic expression of optix in

antennal discs leads to ectopic red pigment in adult antennae (photo courtesy of Makiko Seimiya and Walter Gehring). B: Ectopic expression of eyeless in wing discs leads to ectopic eyes [11] (photo courtesy of Georg Halder). C: The pattern of gene connections between several top regulatory genes in the eye network.

One possibility involves the co-option of the master regulator into a novel context, but at a different expression level from the endogenous eye context, leading to the activation of only those sub-networks that require lower levels of master gene expression to pass an activation threshold.

Previous work characterized the red pigments in Heliconius wings as ommochromes, and proposed that the ommochrome biosynthetic pathway originally deployed in the eye, had been co-opted to the wing. It appears that this was indeed the case, but the co-option was not done in a stepwise fashion. Instead a single gene, that is known to regulate all the elements of this pathway as well as other eye components, optix, was co-opted to the wing.

Diptera из
Балтийского янтаря).
Коопция гена optix в формирование красных

пятен на крыльях Heliconius

Monteiro,​ 2011

строений в результате адаптации к одинаковым условиям.
Концепция адаптивной

зоны Дж. Симпсона: адаптивная зона — комплекс условий внешней среды, определяющая тип адаптаций группы организмов.

George Gaylord Simpson
(1902 — 1984)